Unfortunately, "press 'v' to talk" didn't work, getting me stuck in the first level. :/
(Also, don't forget a way to quit the game, as Alt+f4 shouldn't be relied on)
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You are right that it works while running, which is very odd. Then after the dialogue the game doesn't seem to progress, until then there appears another dialogue option near a random wall to close the "door." Then the boss has some dialogue and the room resets. Also the jump only triggers, on release of space. This is unfortunately too much jank for me to deal with. The art looks amazing.