So much potential here! I really like the look of the characters and abilities. The different abilities themselves are also fun to play with. Really tight controls and the combat hits feel good.
I have similar notes to some others: the NPC convos got in the way sometimes and it's a bummer that you didn't have a chance to set up the unlock progression (though, if the only reason was that you couldn't set up the story, I might've just put the things where you'd wanted them to be anyway, at least then you'd have a game, not just a sandbox).
Having Meg just have a cooldown seems easily spammable by simply waiting a minutes then blitzing through a section, waiting a minute, doing the next section, etc. I might consider having it recharge based on item pick-ups, or only at specific locations (like a recharge booth), or have zones where she's able to activate because of the loose energy in those areas or something.
I can tell there was a lot of work put into this. It's a shame you couldn't quite put it together into a game, woulda been super fun!