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(+2)

The cutscene art is incredible! Impressive pixel art to match. I gave the Forest and Desert bosses a shot, pretty tough stuff! It was somewhat disorienting with both melee and projectile attacks based on the direction you're facing rather than where your cursor is in relation to the player. I think it may be the mouse control. If all of the controls were on the keyboard, the directional attacks would feel more natural. Many boss attacks between the two I played feel sudden and unpredictable, so consider making the telegraphs more apparent or doing some sort of "training" pattern before going into the larger waves of bullets. All in all, good job!

(+1)

Thank you for the art, that was a lot of work indeed, really happy that it seem to please the eye!
I agree the mouse can be disturbing, however it can be played with keyboard only aswell!!
Don't hesitate to give it a try if you want with keyboard only, you can use WASD + IOKL for projectile/shield/dash/attack... (or even better if you have a gamepad controller it works too)
I see what you mean for the telegraph, some patterns are faster than others, which might hit you often, but it should be pretty low damage usually, so it felt like 'necessary' damages to remember the player to not just stay in melee range, and anticipate the potential attacks.  But it would be a great idea to put only the low speed patterns at start to make a sort of tutorial indeed, thank you for the suggestion, it will definitely go in a future update. Also the window of telegraph for most attacks and waves is usually 0.5s to give the player some time to react, but while it seem to be the case for the treant which goes into defensive stance before releasing any pattern, it seem to be less for the sand giant.. Ill dig into it to check. 
Thank you for testing it, and for your kind review!