Wow what a game! First off, the art of the game is just divine(pun intended). Strong Christian symbolism depicted in a wonderful style. Christian/Biblical mythology is mostly used in Japanese games interestingly there are very few western developers that i know which use this theme. Blasphemous series is one of the recent examples. I also know of Dante's Inferno. Btw i'm from Istanbul. In Hagia Sophia there's huge mosaic of Seraphim. I have seen it many times. I used to see it in my nightmares. It's like as if God is watching you with his eye.
Boss design is visually perfect and combat is also very fun and challenging. I like their attack patterns and phases.
Perfect use of the theme in my opinion. You gotta choose between a buff and more time to beat the game. Great execution of risk/reward.
I broke one star from controls but the problem isn't with the controls actually, it's a bug. When you get hit , the frame rate drops to 0 for a seconds than goes back to normal. Fps drop also occurs when you hit the third boss. Sometimes you get hit because the game lags. There's probably a problem in your code, you need to refactor/optimize it.
Music is also perfect, Catholic hymns what else could it be! :) Sfx are nice.
Overall, this is one the best games i've played in the jam so far. I have played about 20 games i think, maybe more. I think this game should be on Top 10 or at least in top 50 but that's not up to me and i haven't seen all the games in the jam yet. I admire your hardwork and creativity, this is a great achievment. I wish you the best of luck in your journey!