I really enjoyed playing the game, when it worked. Unfortunately the game is pretty buggy. Enemies getting stuck in foliage on a massive map is pretty annoying. It's also frustrating that the bases are so far apart. I think I managed to get to the end, but the antagonist didn't die. In the screenshot below the evil controller has zero health. However, the game doesn't say "You Win" or something.
I also noticed that your towers aim at the current position of enemies, resulting in them often missing. You've compensated for this by cranking the firerate up to eleven. For the jam I've also made a tower defence game with a twist. My towers do aim ahead of enemies to account for the time it takes for the projectiles to travel to the future position of the enemies. However, this did bring allong its own issues XD.