I liked the attack variety in the boss fights. Fits the theme with the attack types you chose. Art assets chosen look nice, and the SFX feel appropriate.
The tutorial was very clear and helpful.
For improvements, I feel the melee attack needs improving. I could not work out how to use it effectively. Also the hitbox on the Tiger's Bear Traps is a bit harsh when the player is underneath it, seems to hit the players head a lot
Overall, cool submission, just tighten some of the combat. Nice job!