The story, music and art are uniquely great but I couldn’t get past the first boss. I won’t reiterate some of the problems mentioned but I do have some suggestions that may enhance gameplay and feel. I think the dash and hurt animation are too similar, maybe use a different color for either. I think the boss should have a few more frames of anticipation on the dash attack, adding sound will help as well. Lastly, if you slightly slow down the turret when on the boss collider and speed it up on consecutive misses the player may be able to land more shots.
The color scheme is really cool and a creative take on theme. Good job and keep it up! If I have some time I’ll go back and try to get to the other bosses.