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Do you mean the executable game itself or the source is > 2 GB?

I had the same situation 2 days ago with my source.
My code was stored locally in SourceTree at the time.

I did the following:
Created a GitHub account.
Learnt how to create a remote.
Uploaded to GitHub.
Put the link to the GitHub project in my submission.

If its the game itself my off the top of the head suggestion is:
- make sure you have a backup/source control
- go through the free assetpacks and remove everything you are not using
- rebuild

I'm not sure if UE has the same dynamic resource binding as Unity *shrugs*

Both! 

The textures that my assets are using are 20MB per single file at least. Multiply by 3 (at least) per asset (normals texture, the _D, plus another) times a gazillion assets (lol).... I've dumped everything I'm not using. The packaged project is over 2GB. Can I still upload to itch.io? I'm getting rid of some very heavy foliage assets and replacing with some really low rez ones, but I have an entire interior scene from the dungeon asset pack and its textures are 1.5GB....

I'm trying this setting when UE4 cooks the project with compression enabled. Hopefully that might be better.

But back to itch.io, what's their limit?

Thanks for taking the time to reply! I'll check out your game

Small update ;)

After replacing a very large nature asset pack with some really fuzzy-detail meshes, and enabling compression when packaging, rebuilding the lightmap for my map and enabling this compression option, Unreal's editor packaged it in under 1GB! So, hopefully I'll be ok... Still a rather large submission, though. I don't understand how itch.io's play via browser thing works exactly but I hope players will have no issues trying out the game.

I'm happy to give it a test in case there are issues.
I'm very new here so I don't know what the limits etc really are..