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This happened to me twice as well. The second time I saw that it happened when I reassigned an "idle" villager to destroy a wall. Since the game often reads hunters as idle, it assigned a villager with food in their hands to destroy the wall. The villager was headed to the town center with the food- instead of depositing the food, the villager destroyed the town center. It's game breaking, as doggg91 says. 

Thanks for the detailed report, managed to fix it in v1.4. Download the game again to update! Unfortunately, the saves without a town center will remain without it.

Thanks for the update. Extremely luckily, I rage quit as soon as I saw it happen that second time and when I opened again, the destruction hadn't been completed so I got to finish out that save.