Well done! That's a pretty interesting and challenging metroidvania!
The controls are (as expected) part of the challenge, though I've only tried keyboard and mouse so far. I might give it a shot with a gamepad to see if I can have a better control (I found doing multiple suck in-shoot cycles with jellies to jump over a big gap quite challenging with the mouse, especially since the shoot angle moves at 45° and this can change while I'm jumping if I don't move my mouse properly).
Kudos on all the artistic aspects, it's a pretty good job for a solo game! And it's nice to see everything is properly licensed :)