Another highlight of the jam for me. This is one of the most well put together games here, and as a theme with games of this caliber, so lacking any dramatically unfinished parts, I can only really critique the design.
Fall damage is a little aggressive, since I (like some others) fell prey to fall damage in the blood rune puzzle. The smallest enemies didn't need to both be invisible and leaping, since either one of those things is threatening in the context of your game. Both together makes them seem more tedious than challenging.
Either way! This was one of the most Metroidvania games on the metroidvania game jam. Well scoped and with just enough graphical flourish for it to be exciting to see what else the world had in store!