Wow! This is an incredibly impressive game with amazing art and a great scope. To avoid spoilers, the ending was very impressive! I would say that some really nice music under the sound effects would be nice, but the sound effects themselves are great! Also, the main bug enemies are pretty hard to beat (not necessarily a criticism though) and the camera doesn't always move as far forward as I would like leading to a few spikes...
Overall, a brilliant game that feels right at home with other released indies - if you extended it a bit, you could 100% sell this as a standalone package. Well Done!