The improvements you made to the game over time are pretty impressive ! The world you are creating is very appealing and begs to be expanded. Big plus for the cutscenes and the audio are great, very polished stuff. Congrats on this !
Maybe if had one point to tweek a bit it would be the overall speed of the main character, on foot, on air and while climbing. Slight augmentation could add some dynamics and wouldn't be too off knowing how fast can be the bees at times :)
Keep it up !