Super solid game! The amount of mechanics you guys were able to cram and here are amazing! I also really dig the art a lot. I went for a similar 1-bit with a little extra color splashed in aesthetic on my game but you guys pulled it off way better than me. Also really liked the attention to detail in the design of the camera. It was cool how it focused on different areas depending on which area you were in. Did you guys do this with cinemachine? I unfortunately got stuck in the level that introduced swimming and couldn't figure out how to finish it. I wouldn't take this as a knock to your level design though. Even though I love metroidvanias I get stuck in them very often even in ones made by AAA teams.
One very small critique. I think overall the sfx were incredible and especially liked the walking sound one. But I found it slightly odd that the walking sound was heavily reverbed and the jump sound was completely dry. Also the jump sound seemed to cancel out the reverb tail on the walking sound. If you use playoneshot instead of play it shouldn't do this. Either way obviously a super minute critique on any game especially a jam game, but I figured it would be super quick to change if you guys decide to work on the game further and it in no way will effect my rating. Really good job on your game!