Like others have said, the art style is really great and definitely stands out. I appreciate how you re-used a lot of the pre-existing game mechanics in the boss fight (It would have been cool if all three gameplay styles came back in some way, although I'm not sure how you'd incorporate the runner section- which is probably why you didn't lol)
I think the game gets a little hard to read during the boss fight. I kept taking damage from bugs I didn't really notice and walking off ledges trying to line myself up for the block puzzle.