That was really cool! I totally didn't expect all of that. I love how both the puzzle and action parts were both very well-made and had a lot of thought put into them. The story is hilarious and I think you are absolutely great at writing dialogues.
However, if it weren't for the amazing graphics and music, I totally wouldn't like to play longer than few screens at first, because there was no introduction or explanation what I'm supposed to do. Also, I've seen a lot of people complaining about the last boss being hard, I just think that was really annoying and definitely the worst part of the game for me. But it's an easy fix, just give him a bigger hitbox and make the mole's movement more controllable. I understand there wasn't probably enough time for polishing like that, but I think the game is really professional and you could have people playing it outside of the jam, so think about those improvements.
You did an amazing job :)