This was really fun. The introductions of new mechanics were well done and I liked the fast-paced aiming + running through a lot of the levels. Polish is there, sound effects work well, and the visual style, though simplistic, does its job well and remains cohesive.
My only criticism is that the time limit didn't seem as justified in the more puzzle-focused stages. A few, as simple as they were, really wanted me to stay still and survey the area before continuing forward. (Maybe you could have a bomb item or something specifically for fast-paced sections?)