Really great atmosphere, Good tutorial(Maybe missing add lock-on distance explanation). The mechanics work great, the music is great and I loved seeing the boss move-set and finding the strategy to defeat him.
For improvement, I would say the controls felt pretty "clunky" and the spawn of the enemies in the last room made it so every time they spawn they would shoot at the same moment or spawn on the player etc.
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I actually have a solution for the spawning problems including the ones you didn't mention. But I don't know how to deal with the "clunky" controls. Maybe it's because the animations are too slow, so it feels like some old RPG games out there. I actually feels the same if you're talking about this exact thing. I'd need to find a good way to find the right pace for the game. As for the target-lock distance....well I actually have no idea how to explain it. Maybe make indicator to show if the enemies are within the range, I think this should be good enough.