Fun little game! I appreciated how you incorporated the theme into the mechanics. It was also good to see a tutorial level, key-rebinding, and some accessibility options. I think the levels scaled in difficulty reasonably well, and I found the ghost level a fun way to subvert some of my expectations.
I preferred the shorter levels that reduced the punishment for failure. I think impatient players would appreciate a checkpoint in the last level, or splitting it into two. Some of the default control choices went against conventions (C for run), so it was good to offer re-binding. Though the rebinding captures mouse motion as an input, which is probably not intended.
A very solid first jam submission. I look forward to your next game!