I love roguelikes and kudos to you for making one for a game jam. The shooting mechanic is very interesting and unique, however it wasn't my cup of tea. I found that the best way to proceed is to just kite the enemies into a big bunch and then shoot them, and this became very repetitive very fast. The upgrade system is cool, but it kinda feels like the Shrine theme was shoehorned into the game, like the upgrade shrines could have been anything else and it wouldn't have changed the feel of the game much.
Also not sure if this is intended but sometimes I would take damage from the spikes even when I am running across them, felt like a game of chance whether or not I would take random damage form crossing spikes.