I really liked the concept and looks a lot, crunchy textures for the win! Probably my favorite boss was the one at the river because I used the harpoon gun and got to be constantly spining around jaja.
In terms of optimization it was a hit or miss depending on the bossfight (mostly played in browser), the snake arena in therms of design for me was a bit rough specially needing to look up almost constantly and damage or ammo of weapons could have been a bit high in general. Other than that this was a banger, would pay for a full release no doubt!!