Playing it was fun; actually winning was unpleasant. I find the difficulty completely inscrutable; I actually downloaded the windows build to make sure I wasn't getting weird web errors.
I rarely ask to be spoon-fed information, but I had no clue how health works. There were several times when I died when I thought I had only been hit twice, making me wonder where I actually lost all that health. I was also never sure how much damage each specific tower actually dealt. I'm not necessarily suggesting a literal health bar, just more specific feedback on damage and health.
Volume settings are appreciated, but the default audio volume is a tiny smidgen loud; later parts sometimes peak above +0.0 dB.
All-in-all, good work!