The game feels very nice and I support the core idea with my full passion. Portraying humans as overexpanding threat is what games rarely do, but this is a very important message.
The music fits the theme and tempo of the game pretty good. Art looks coherent and beautiful.
As for the game mechanics — it looks a little bit too random, as if the game lacks ways to communicate to the player when and where humanity will expand. I lost some games placing just a few tiles, and won some just randomly dropping the stuff on board.
The overall logic seems understandable — mountains can cut off cities, cities would grow to adjacent tiles (sometimes?), rivers are op but you need to roll for them before tiles are getting used for other stuff etc. But I found it hard to actually strategically plan on what to do, as if random tiles are more responsible for me winning or not.
I think that two main point on how to improve fun of the game are:
1. More clear communication on "human turn". When they will expand and where, so the player can act in advance and stop it.
2. Less influence of random. To make more options in placing tiles and their types. Maybe simply expanding the number of available tiles from 2 to 3 will do, or changing the logic from total random to include some important types of tiles once in a while.
Anyways cool concept and a very much playable prototype. Good work!