One little nitpick I have with the writing is that when Asterion, P or Storm (IIRC) are anxious they're often feeling "Hummingbirds" pecking on their neck or spine.
I started to notice this metaphor crop up again and again until it became almost a personal meme, which made me feel that some of the instances could be replaced with something else.
If there is any thematic significance to the hummingbirds I didn't notice it, it's not something someone would say where I am from (butterflies in the stomach?) but that may be because there are no native species of hummingbirds here :)
Because there are anthro characters in this world any animal metaphor does also run the risk of being taken literally, could there be a sinister hummingbird mythical plotting their demise?
In all seriousness it's a cute metaphor but I would personally reduce how often it crops up, or make it so only one character experiences anxiety in this way, unless the purpose was to show their parallel journeys?