This game is Neat ;)

• It feels very fleshed out with lots of content; your 2 years of work definitely shows in the game
• Beautiful graphics with soothing color schemes
• Fun bosses with interesting mechanics
• Interesting world that makes me want to keep exploring
• The difficulty feels good (coming from someone who doesn't play metroidvania's mind you)

A couple humble suggestions:

• Remappable buttons (I used an Xbox controller and it felt unconventional to press Y for interact and B for melee)
• Ability to re-initiate dialog (in case you forgot what the NPC said)
• The beetle is a cool enemy; it would add a little interest if it could only be damaged from the rear since its face looks pretty tough and armored
• The graphics are beautiful; I understand the aesthetic you're going for, but perhaps adding more contrast to a small number of elements may help it pop out in the screenshots (this is assuming you're going for a commercial release)
• A quality of life feature would be to have the interaction button hints match your current control scheme (when I play with gamepad the hints are for the keyboard keys). I saw a tutorial on this the other day: 

Talk about polish, this game honestly feels commercial release ready. The soundscapes are nice, the art looks good, the controls are well laid out and feel good. The game as a whole is fun to play and feels smooth, great work!

This game is very polished, the mechanic is cool and everything works perfectly. I would not call it "pre-alpha" :)

It is not my genre (I never played another metriodvania - had to look it up on wikipedia) so it is hard for me to give you any advice. There are no obvious shortcomings ...

What I liked most where the boss fights - figuring out the mechanic and then applying the "win"-strategy was engaging.

Unfortunately I'm on a Mac, so I'm not able to do the Windows download. But I just wanted to comment to say that I remember seeing posts about your game on Reddit a while back, so the mechanic was definitely interesting enough to me that I remember it after all this time! Looks like an interesting and fun concept!

Honestly, your game is good. I think the biggest issue you're going to face as far as a commercial release goes is that even if your game is good, the competition in the platformer/metroidvania genre is the highest bar possible.

That being said, I do think you've cooked up something quite nice here. The actual mechanics feel really good and exploring is fun. The enemy designs were interesting too. If there was anything I had to say was wrong with it, it'd have to be the protagonist not really looking interesting. I didn't play very far (I killed the spitting beetle boss guy and got acorn mode) but I hope there's unique biomes later in the game too.

See it through to the end! Good luck!

I was just about to ask you how much time you spent on it and 2 years is crazy! It's an amazing game very smooth. Amazing gameplay and the dialogue system too. Did you use any libraries to help you speed up development?